Earlier quoted context omitted.
> "cheating" and infidelity are largely human concepts It is normal behavior, yes. But it comes at the expense of the cheated on party. The term here is "parental investment" and is roughly how much energy a party has to invest in the offspring. If the male tit builds the nest and provides food for offspring that is genetically not his it puts it at a disadvantage and it is definitely exploited by the female.
Again, "exploited"? Using that word in the context seems very... Off. It's not like the female tit is choosing to "exploit" the male. They're literally programmed to do that. Think about it. In many species there is unequal parental employment. That's just how nature works. Is it really "exploitation" in the human sense of taking advantage of someone else?
Yes. Obviously. And whether or not the exploitation is "programmed" is irrelevant. After all everything is programmed.