Earlier quoted context omitted.
The whole point of the article is that the traditional "calories in/calories out" model is wrong. "Based on the research, Pontzer has proposed a new model that upends the the old "calories in, calories out" approach to exercise... He calls this the "constrained model" of energy expenditure, which shows that the effect of more physical activity on the human body is not linear."
The conclusion is: "10) So what actually works for weight loss? At the individual level, some very good research on what works for weight loss comes from the National Weight Control Registry, a study that has parsed the traits, habits, and behaviors of adults who have lost at least 30 pounds and kept it off for a minimum of one year. They currently have more than 10,000 members enrolled in the study, and these folks…
The traditional model proposes a much simpler linear relationship: do an additional unit of exercise and you will burn X more calories, forever. He says that is incorrect and you actually get diminishing returns, where additional units of exercise burn ever fewer calories rather than just a constant.