The author's examples are pretty horrible, but consider... Xobni is an extension to outlook, and yet they have built a solid business from this.
Flickr's photo sharing was a feature of a game they were developing before they turned it into a stand-alone business.
Blue-Ray, Dolby, THX can all be considered 'features' as they are not a product in themselves, but an improvement of an existing product. Gracenote & AMG metadata services.
I suspect many of these were originally thought of as features before a business model was discovered for them.