also noteworthy elements: university overhead 53%, so that leaves $250K. Over five years that works out to basically $50K/year That needs to cover 1 month summer salary plus the salary and the tuition for a single student, and the travel to conferences, and the experiment ... and that is a "prestigious" CAREER award only few people get ... no wonder the author jumped ship
So for a $500K grant, 53% overhead really means about 34.6% of $500K. The university's cut is $173K, which is 53% of the PI's budget of $327K.