From the article:

The critics argue that Nowak and his colleagues make an unwarranted distinction between inclusive fitness and natural selection theory, and assert that inclusive fitness has been quite valuable. They stress that inclusive fitness ideas have helped yield insights into many biological phenomena, such as why social insects have skewed sex ratios. "Their announcement of the demise of inclusive fitness is wrong," says STRI's William Wcislo. "They've done a service in a sense: [Their paper] forced people to again rethink something, and it's quite clear that the evidence in favor of Hamilton's theory is quite convincing."