So to recap, Twitter exploded onto the scene in 2007, the "fail whale" appeared a lot, developers made all sorts of wonderful programs hooked into Twitter, the fail whale disappeared, Twitter started to destroy the app ecosystem, App.net launched to great fanfare in response to Twitter's knuckleheaded anti-developer stance, Britney Spears and Justin Bieber arrived and knocked all the nerds out of the top spots on Twi…
Twitter finances (TTM): revenue: $2.52B gross profit: $1.49B It's only losing money because of stock grants.
TL;DR -- Twitter paid out about $680M in grants