Earlier quoted context omitted.
I think you just answered your own question. I'd say just about every Unix box on the planet has Perl holding it together somewhere in there. What you've asked is essentially like asking "How does mortar hold buildings together any more than bricks, steel I-beams, or roofing?" These things are equally important, and yes you could probably find a substitute, but why?
"What you've asked is essentially like asking "How does mortar hold buildings together any more than bricks, steel I-beams, or roofing?" Argument by metaphor is a lazy device. I don't see any specific "mortar like" virtues that perl has but other common scripting languages don't. In this metaphor I guess C Code would be the "bricks and steel beams". Why can't I put a unix box together without perl on it? Is there any…
It means that Perl is used in a lot of unexpected places, and that a lot of critical things are "held together" by Perl. Even if you think your website is running on Java, some systems administrator probably has a cronjob written in Perl critical to its operation.