With so many accounts compromised, the hackers might actually have full access to Twitter's backend. The postmortem would be very interesting. I'll be looking forward to it. Imagine if the hackers timed the intrusion during github outage, and twitter's employees can't deploy a fix for the exploit fast enough because github was down!
It seems like the devs at Twitter are clueless, how this happened. The hackers could be deep in Twitters systems, eventually even have even someone working at Twitter, or it's a result of a new yet unknown password list or phishing attempt.
Means they had someone inside Twitter.