I feel like the paper lacks a discussion about why males are more likely to take risks. Typically extremely reproductively successful men like Chengis Khan take enormous risks to get the power and status that leads to high reproductive success. Unfortunately we are all descended from those men.
I don't feel like there needs to be a discussion on 'why' they take more risks. Females expand way more energy in having and raising kids than males do (within the human species). In other species (like sea-horses) - the females actually compete for males since males are the ones that expand way more energy in raising / producing children... (i.e. https://oceanservice.noaa.gov/facts/seahorse.html ).
In my first year at college we were all set an essay about why human females invest lots of resources preening and enhancing their appearance, while across nature females normally put almost no effort into their appearance, and males are the ones with the bright colours.
This is an enormous area of study with thousands of papers published. Sexual selection is incredibly important. Especially as modern human evolution is driven almost entirely by sexual selection.
It’s worth understanding how sexual selection works.