Reproductive analogy:
A sequence of AI models trained on each other's output gets mutations, which might help or hurt, but if there's one dominant model at any given time then it's like asexual reproduction with only living descendant in each generation (and all the competing models being failures to reproduce). A photocopy of a photocopy of a photocopy — this seems to me to also be the incorrect model which Intelligent Design proponents seem to mistakenly think is how evolution is supposed to work.
A huge number of competing models that never rise to dominance would be more like plants spreading pollen in the wind.
A huge number of AI there are each smart enough to decide what to include in its training set would be more like animal reproduction. The fittest memes survive.
Memetic mode collapses still happen in individual AI (they still happen in humans, we're not magic), but that manifests as certain AI ceasing to be useful and others replacing them economically.
A few mega-minds is a memetic monoculture, fragile in all the same ways as a biological monoculture.