Speaking as someone who lived through this period during the formative years of my programming career, the author has completely missed the most influential language of the time and, in my mind at least, the number reason why Perl lost. Yes, I'm talking about Java. It's easy to hate now, but Java back then replaced all the server-side Perl programming that I did in the space of about 3 years, from 1994 where CGI prog…
There's a wrinkle, too. Perl isn't just a web thing. It was (and is) still used in "enterprise" spaces for scripting on servers. It's still out there being used to create scripts rather than applications. In that space Ruby/Python are competing, but there's still plenty of sysadmins who use Perl since it works.